John W. Burch, National Park Service, 201 1st Avenue,
Fairbanks, AK 99701, USA; Layne G. Adams, U.S. Geological Survey,
Biological Resources Division, Alaska Biological Science Center,
1011 East Tudor Road, Anchorage, AK 99503, USA; Erich H. Follmann,
Eric A. Rexstad, Institute of Arctic Biology, University of
Alaska Fairbanks, Fairbanks, AK 99775, USA
Density estimation of wolves (Canis lupus) requires
a count of individuals and an estimate of the area the wolf
population inhabits. With radiomarked wolves, the count of wolves
is straight forward but the estimation of area is more difficult
and often not given adequate attention. The population area,
based on the mosaic of pack territories, is influenced by sampling
intensity in a similar manner as individual home ranges. If
sampling intensity is low, the population area will be underestimated
and wolf density estimates will be inflated. Using data from
studies in Denali National Park, Alaska, we investigated these
relationships using Monte Carlo simulation to evaluate the effects
of radiolocation effort and the number of marked packs on density
estimation.
We report that as the number of adjoining pack home ranges
increase, fewer locations are necessary to define a given percentage
of population area. We evaluated the utility of nonlinear regression
to adjust for biases associated with under sampling and present
recommendations for monitoring wolves via radiotelemetry.